S = cAz — power-law biodiversity and the fragmentation cost to ecosystems
SAR Parameters
Fragmentation
Predictions
Species (intact A₀)—
Species (per fragment)—
Total species (fragments)—
Species lost—
% biodiversity lost—
z typically 0.25–0.30 for mainland. Fragmentation: same total area, but split into N patches each with area A₀/N, losing species to edge effects and isolation.