Active strategies:
PD condition: T > R > P > S (and 2R > T+S). In a well-mixed population,
Defection dominates. Tit-for-Tat (Axelrod 1984 tournaments): cooperate first, then copy opponent.
Spatial structure (Nowak & May 1992): cooperators cluster, resist defector invasion.
Win-Stay Lose-Shift outperforms TFT in noisy environments.
Evolution of cooperation requires: kin, reciprocity, network, group, or reputation mechanisms.