Activator-Substrate Depletion — Reaction-Diffusion Patterns

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Activator-substrate (Gierer-Meinhardt substrate-depletion) model: dA/dt = ρA²/S − κA + DaΔA; dS/dt = ρ₀ − ρA² + DsΔS. Unlike activator-inhibitor, here the substrate is consumed, creating depletion zones. Produces spots, stripes, and labyrinthine patterns.