Replicator-Mutator Eq.

ẋᵢ = Σⱼ Qⱼᵢ fⱼ xⱼ − φ xᵢ
Mutation matrix Q couples evolution to error correction. Models viral quasispecies & language change.

Mean fitness φ:
Diversity H:
Error threshold: μ* ≈ ln(s)/L