Ecological Niche & Competitive Coexistence

MacArthur's consumer-resource model: niche overlap, exclusion, and neutral theory

MacArthur-Wilson Niche Theory

Each species has a Gaussian utilization function u_i(r) centered on its optimal resource. The competition coefficient α_ij = ∫u_i u_j dr measures niche overlap.

Competitive exclusion principle (Gause 1934): two species sharing a single resource cannot stably coexist. With many resources, coexistence requires niche differentiation — niche width σ must be small enough that overlap α_ij ≪ 1.

Neutral theory (Hubbell 2001): small immigration m prevents extinction and maintains diversity even without niche differences. The Unified Neutral Theory predicts log-normal species abundance distributions.

MacArthur consumer-resource competitive exclusion neutral theory