Chromatin Remodeling

ATP-dependent nucleosome sliding by SWI/SNF and ISWI complexes
Remodeling Controls
ATP Budget
ATP: 100%
Nucleosomes: 3
Open bp: 0
Mode: Idle
Biology
SWI/SNF: ~1 MDa complex
Smarca4/BRG1 ATPase
Slides or ejects nucleosomes

ISWI (NURF/CHRAC):
Spaces nucleosomes evenly
Linker DNA sensing by AutoN

ATP hydrolysis pumps
DNA around the octamer
~10-25 bp/ATP