Adaptive Dynamics & Evolutionary Branching

Invasion fitness, PIP plots, and evolutionary speciation

Pairwise Invasibility Plot (PIP)

Evolutionary Dynamics

Invasion Fitness

Theory

f(y,x) = r[1 − (α(y−x)·n*)/K(y)]
K(x) = K₀ exp(−x²/2σ_K²)
α(x) = exp(−x²/2σ²)
ẋ = μ ∂f/∂y|_{y=x}

PIP plot: green = mutant y can invade resident x; red = cannot. Singular strategy x* where ∂f/∂y=0. If also ∂²f/∂y²>0 (convergence stable) and locally uninvadable → ESS. If convergence stable but invadable → evolutionary branching point.


When σ < σ_K, the singular point is a branching point: after convergence, a rare mutant can invade and the population splits into two distinct phenotypes — evolutionary speciation.